COURSES
offered in the PhD Program in Biology
General
Information
60000-level graduate courses are listed in the graduate bulletins
of the City, Hunter, Brooklyn, Queens, and Lehman Colleges of
the University. A maximum of 9 credits in such courses may be
offered toward the Ph.D. degree.
70000- and 80000-level graduate courses (University courses)
creditable toward the doctoral degree are listed below. Each
course is offered periodically at one or more units of the University
as indicated. Courses listed under the same University number
cover substantially similar material at comparable levels. For
course descriptions consult the several graduate bulletins of
each college of the University. The prerequisite for admission
to all courses is prior approval by the student's advisory committee.
The specific offerings at The Graduate Center and each campus
are listed in the Announcement of Courses issued prior to each
semester.
For classes in a specific subprogram, select one of the
links below:
Ecology,
Evolutionary Biology, and Behavior
Molecular,
Cellular, and Developmental Biology
Neuroscience
Plant
Sciences
Seminars:
General and Special Topics
General
Ecology,
Evolutionary Biology, and Behavior
BIOL 70003 Genetics: Lecture; 30 or 45 hours lecture,
2 or 3 credits
BIOL 70004 Genetics: Laboratory; 60 or 90 hours laboratory,
2 or 3 credits
BIOL 70005 Genetics: Lecture; 60 or 75 hours lecture,
4 or 5 credits
Prerequisite: Undergraduate genetics and molecular biology
or biochemistry
BIOL 70006 Genetics: Lecture; 45 hours lecture, 3 credits
BIOL 70103 Microbial Genetics; 45 hours lecture, 3 credits
BIOL 70104 Problems in Microbial Genetics: Laboratory; 90
hours laboratory, 3 credits
BIOL 70201 Genetics of Multicellular Organisms, I (H); 45
hours lecture, 3 credits
BIOL 70202 Genetics of Multicellular Organisms, II; 45
hours lecture, 3 credits each semester
BIOL 70301, 70302 Cell Heredity I, II; 45 hours lecture,
3 credits each semester
BIOL 70503 Evolution; 30 or 45 hours lecture, 2 or 3
credits
BIOL 70504 Evolution: Laboratory; 60 hours laboratory,
2 credits
BIOL 70505 The Evolutionary Biology of Vertebrates; 45
hours lecture, 3 credits
BIOL 70506 Macroevolution: Patterns of Evolution above the
Species Level; 45 hours, 3 credits
BIOL 70603 Principles of Systematics; 45 hours lecture
plus conferences, 4 credits
BIOL 70604 Animal Systematics: Laboratory; 60 or 90 hours
laboratory, 2 or 3 credits
BIOL 70609 Biological Museology; 45 hours, 3 credits
BIOL 70611 Systematics and Evolution of Insects and Arachnids:
Lecture; 2 hours, 2 credits
BIOL 70612 Systematics and Evolution of Insects and Arachnids:
Laboratory; 4 hours, 2 credits
BIOL 70713 Zoology and Phylogeny of Chordata (Fishes); 30
hours lecture, 2 credits
BIOL 70714 Zoology and Phylogeny of Chordata (Fishes); 60
hours laboratory, 2 credits
BIOL 70723 Zoology and Phylogeny of Chordata (Reptiles and
Amphibians); 30 hours lecture, 2 credits
BIOL 70724 Zoology and Phylogeny of Chordata (Reptiles and
Amphibians); 60 hours laboratory, 2 credits
BIOL 70733 Zoology and Phylogeny of Chordata (Mammals); 30
hours lecture, 2 credits
BIOL 70734 Zoology and Phylogeny of Chordata (Mammals); 60
hours laboratory, 2 credits
BIOL 70735 Functional and Adaptational Biology of the Mammalia; 45
hours, 3 credits
Prerequisite: 70733, 70734
BIOL 70743 Zoology and Phylogeny of Chordata (Birds); 30
hours lecture, 2 credits
BIOL 70744 Zoology and Phylogeny of Chordata (Birds); 60
hours laboratory, 2 credits
BIOL 70800 Biochemical Evolution and Systematics; 45
hours lecture, 3 credits
BIOL 70803 Molecular Evolution; 45 hours, 3 credits
BIOL 70901 Population Genetics: Lecture; 45 hours lecture,
3 credits
BIOL 70902 Population Genetics: Laboratory; 90 hours
laboratory, 3 credits
BIOL 70903 Quantitative Genetics; 45 hours lecture, 3
credits
BIOL 70907 Behavior Genetics; 30 hours lecture, 120 hours
laboratory, 4 credits
BIOL 72403 Animal Behavior I: Lecture; 45 hours lecture,
3 credits
BIOL 72404 Biological Basis of Animal Behavior: Laboratory; 90
hours laboratory, 3 credits
BIOL 72405 Current Issues in Behavioral Ontogeny; 45
hours lecture, 3 credits
BIOL 72406 Behavior and Evolution; 45 hours lecture,
3 credits
BIOL 72407 Animal Behavior II; 45 hours, 3 credits
BIOL 72505 Animal Communication: Lecture; 45 hours lecture,
3 credits
BIOL 72800 Field Studies in Animal Behavior; 90 hours
fieldwork and conferences, 3 credits
BIOL 73103 Microbial Ecology: Lecture; 30 hours lecture,
2 credits
BIOL 73104 Microbial Ecology: Laboratory; 90 hours laboratory,
3 credits
BIOL 74103 Radiation Biology: Lecture; 30 or 45 hours
lecture, 2 or 3 credits
BIOL 74104 Radiation Biology: Laboratory; 60 or 90 hours
laboratory, 2 or 3 credits
BIOL 76001 Ecology: Lecture; 45 hours lecture, 3 credits
BIOL 76002 Ecology: Laboratory and Field Study; 90 hours
laboratory, 3 credits
BIOL 76003 Community Ecology: Lecture; 45 hours lecture,
3 credits
BIOL 76004 Community Ecology: Laboratory; 90 hours laboratory,
3 credits
BIOL 76005 Population Ecology: Lecture; 45 hours lecture,
3 credits
BIOL 76006 Population Ecology: Laboratory; 90 hours laboratory,
3 credits
BIOL 76007 Limnology: Lecture; 45 hours lecture, 3 credits
BIOL 76008 Limnology: Laboratory; 90 hours laboratory,
3 credits
BIOL 76101 Marine Plankton Dynamics: Lecture; 45 hours
lecture, 3 credits
BIOL 76102 Marine Plankton Dynamics: Laboratory; 90 hours
laboratory and field trips, 3 credits
BIOL 76103 Marine Benthos: Lecture; 45 hours lecture,
3 credits
BIOL 76104 Marine Benthos: Laboratory; 90 hours laboratory
and field trips, 3 credits
BIOL 76105 Fishes and Fisheries Biology: Lecture; 45
hours lecture, 3 credits
BIOL 76106 Fishes and Fisheries Biology: Laboratory; 90
hours laboratory, 3 credits
BIOL 76107 Marine Microbiology: Lecture; 45 hours lecture,
3 credits
BIOL 76108 Marine Microbiology: Laboratory; 90 hours
laboratory and field trips, 3 credits
BIOL 76113 Marine Ecology: Lecture; 30 hours lecture,
2 credits
BIOL 76114 Marine Ecology: Laboratory; 60 hours laboratory,
2 credits
BIOL 76200 Physiological Ecology: Lecture; 45 hours lecture,
3 credits
BIOL 76201 Physiological Ecology: Laboratory; 90 hours
laboratory, 3 credits
BIOL 76701 The Biology of Fishes; 45 hours lecture, 3
credits
BIOL 76702 The Biology of Fishes; 90 hours laboratory,
3 credits
BIOL 76830 World Vegetation; 45 hours, 3 credits
BIOL 77200 Biological Electron Microscopy; 30 hours lecture
or demonstration, 90 hours laboratory, 4 credits
BIOL 78001 Mathematical Biology: Lecture; 30 hours lecture
and conferences, 3 credits
BIOL 78002 Mathematical Biology: Laboratory; 60 or 90
hours laboratory, 2 or 3 credits
BIOL 78101 Advanced Mathematical Biology: Lecture; 30
hours lecture and conferences, 3 credits
BIOL 78102 Advanced Mathematical Biology: Laboratory; 60
or 90 hours laboratory, 2 or 3 credits
BIOL 78201 Biostatistics I; Lecture/Laboratory 3 hours
lecture/6 hours laboratory, 6 credits
BIOL 78202 Biostatistics II; Lecture/Laboratory 3 hours
lecture/6 hours laboratory, 6 credits
BIOL 79001 Colloquium in Ecology, Evolution, and Behavior; 15
hours, 1 credit
BIOL 79030 Seminar in Ecology, Evolution, and Behavior; 15
hours, 1 credit
BIOL 80803 Microevolutionary Processes: Lecture; 30 or
45 hours lecture, 2 or 3 credits
BIOL 80804 Microevolutionary Processes: Laboratory; 60
or 90 hours laboratory, 2 or 3 credits
Top
Molecular, Cellular, and Developmental Biology
BIOL 70400 Problems in Nuclear Cytology; 30 hours lecture,
60 hours laboratory, 4 credits
BIOL 70801 Developmental Genetics: Lecture; 45 hours
lecture, 3 credits
BIOL 70802 Developmental Genetics: Laboratory; 90 hours
laboratory, 3 credits
BIOL 71013 Molecular Biology: Lecture; 75 hours lecture,
5 credits
BIOL 71014 Molecular Biology: Laboratory; 15 hours lecture,
90 hours laboratory, 4 credits
BIOL 71015 Molecular Biology: Lecture; 60 hours, 4 credits
BIOL 71016 Molecular Biology: Laboratory; 90 hours laboratory,
3 credits
BIOL 71101 Laboratory Rotation; 90 hours, 3 credits
BIOL 71103 Experimental Microbiology: Lecture; 45 hours
lecture, 3 credits
BIOL 71104 Experimental Microbiology: Laboratory; 90
hours laboratory, 3 credits
BIOL 71203 Comparative Biochemistry: Lecture; 30 hours
lecture plus conferences or 45 hours lecture; 3 credits
BIOL 71204 Comparative Biochemistry: Laboratory; 60 or
90 hours laboratory, 2 or 3 credits
BIOL 71300 Biology of Aging; 45 hours lecture, 3 credits
BIOL 71401 Cell Biology: Lecture; 60 or 75 hours lecture,
4 or 5 credits
BIOL 71402 Cell Biology: Laboratory; 180 hours laboratory,
6 credits
BIOL 71403 Cell Biology: Lecture; 30 hours lecture, 2
credits
BIOL 71404 Cell Biology: Laboratory; 90 hours laboratory,
3 credits
BIOL 71500 Cell Biology Internship; 20 hours week, literature
assignments and laboratory work, 10 credits
BIOL 71600 Cells in Culture; 60 hours laboratory, 15
hours recitation, 3 credits
BIOL 71700 Bacteriophage; 45 hours lecture, 3 credits
BIOL 71710 Virology; 3 hours lecture, 3 credits
BIOL 71800 Immunology; 45 or 60 hours, 3 or 4 credits
BIOL 71903 Medical Microbiology and Immunology; 67.5
hours lecture, 22.5 hours laboratory, 22.5 hours conference,
6 credits Prerequisite: BIOL 71800 or equivalent and permission
of instructor
BIOL 74000 Introduction to Biophysics; 45 hours lecture,
3 credits
BIOL 74001 Biophysical Techniques in Physiology; 90 hours
laboratory, 3 credits
BIOL 74200 Radioisotopes in Biology; 60 hours lecture,
laboratory and demonstrations, 4 credits
BIOL 74300 Photobiology; 45 hours lecture, 3 credits
BIOL 74400 Electrobiology; 45 hours lecture, 3 credits
BIOL 74700 Structure and Metabolism of Macromolecules; 30
hours lecture, 30 hours laboratory, 3 credits
BIOL 75003 Developmental Biology: Lecture; 45, 60, or
75 hours lecture, 3, 4, or 5 credits
BIOL 75004 Developmental Biology: Laboratory; 60 or 90
hours laboratory, 2 or 3 credits
BIOL 75101 Special Problems in Developmental Biology: Lecture; 30
hours lecture, 2 credits
BIOL 75102 Special Problems in Developmental Biology: Laboratory; 60
hours laboratory, 2 credits
BIOL 75303 Molecular Basis of Development: Lecture; 30
or 45 hours lecture, 2 or 3 credits
BIOL 75304 Molecular Basis of Development: Laboratory; 90
hours laboratory, 3 credits
BIOL 76300 Experimental Parasitology: Lecture; 45 hours
lecture, 3 credits
BIOL 76301 Experimental Parasitology: Laboratory; 90
hours laboratory, 3 credits
BIOL 77003 Cytology: Lecture; 30 or 45 hours lecture,
2 or 3 credits
BIOL 77004 Cytology: Laboratory; 60 or 90 hours laboratory,
2 or 3 credits
BIOL 77100 Problems in Experimental Cytology; 30 hours
lecture, 90 hours laboratory, 5 credits
BIOL 77101 Analysis of Mammalian Cells in Tissue Culture; 30
hours lecture, 90 hours laboratory, 5 credits
BIOL 77301 Cytogenetics: Lecture; 45 hours lecture, 3
credits
BIOL 77302 Cytogenetics: Laboratory; 60 hours laboratory,
2 credits
BIOL 77400 Basic Principles of Cellular Microsurgery and
Micromanipulation; 30 hours lecture, 30 hours laboratory,
3 credits
60 or 90 hours laboratory, 2 or 3 credits
Top
Neuroscience
BIOL 71003 Cellular Physiology: Lecture; 30 or 45 hours
lecture, 2 or 3 credits
BIOL 71004 Cellular Physiology: Laboratory; 60 or 90
hours laboratory, 2 or 3 credits
BIOL 72001 Animal Physiology: Lecture; 45 or 60 hours
lecture, 3 or 4 credits
BIOL 72002 Animal Physiology: Laboratory; 90 hours laboratory,
3 credits
BIOL 72101 Animal Physiology I; 60 hours, 4 credits
BIOL 72102 Animal Physiology II; 60 hours, 4 credits
BIOL 72201 Endocrinology: Lecture; 45 hours lecture,
3 credits
BIOL 72202 Endocrinology: Laboratory; 90 hours laboratory,
3 credits
BIOL 72301 Neurosciences I: Lecture; 45 hours lecture
plus recitation, 4 credits
BIOL 72302 Neurosciences II: Lecture; 45 hours lecture
plus recitation, 4 credits
BIOL 72303 Neurosciences II: Laboratory; 120 hours laboratory,
4 credits
BIOL 72401 Comparative Neuroendocrine Mechanisms; 45
hours lecture, 3 credits
BIOL 72403 Animal Behavior I: Lecture; 45 hours lecture,
3 credits
BIOL 72404 Biological Basis of Animal Behavior: Laboratory; 90
hours laboratory, 3 credits
BIOL 72405 Current Issues in Behavioral Ontogeny; 45
hours lecture, 3 credits
BIOL 72406 Behavior and Evolution; 45 hours lecture,
3 credits
BIOL 72407 Animal Behavior II; 45 hours, 3 credits
BIOL 72503 Sensory Physiology: Lecture; 45 hours lecture,
3 credits
BIOL 72504 Sensory Physiology: Laboratory; 90 hours laboratory,
3 credits
BIOL 72505 Animal Communication: Lecture; 45 hours lecture,
3 credits
BIOL 72603 Comparative Animal Physiology: Lecture; 30
or 45 hours lecture, 2 or 3 credits
BIOL 72604 Comparative Animal Physiology: Laboratory; 60
or 90 hours laboratory, 2 or 3 credits
BIOL 72703 Endocrine Cytology: Lecture; 45 hours lecture,
3 credits
BIOL 72704 Endocrine Cytology: Laboratory; 60 hours laboratory,
2 credits
BIOL 72800 Field Studies in Animal Behavior; 90 hours
fieldwork and conferences, 3 credits
BIOL 72901 Trends and Issues in Physiology/Neuroscience I; 3
hours lecture during the Fall, alternate weeks for 1 credit
by two or three doctoral faculty at The Graduate Center
BIOL 72902 Trends and Issues in Physiology/Neuroscience II; 3
hours lecture during the Spring, alternate weeks for 1 credit
by two or three doctoral faculty at The Graduate Center
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Plant Sciences
BIOL 70601 Plant Systematics: Lecture; 30 hours lecture, 2
credits
BIOL 70602 Plant Systematics: Laboratory; 60 hours laboratory,
2 credits
BIOL 70605 Taxonomy of Vascular Plants: Lecture; 45 hours
lecture, 3 credits
BIOL 70606 Taxonomy of Vascular Plants: Laboratory; 60 hours
laboratory, 2 credits
BIOL 70607 Numerical Systematics: Lecture; 30 hours lecture,
2 credits
BIOL 70608 Numerical Systematics: Laboratory; 60 hours laboratory,
2 credits
BIOL 73001 Plant Physiology: Lecture; 30 or 45 hours lecture,
2 or 3 credits
BIOL 73002 Plant Physiology: Laboratory; 60 or 90 hours laboratory,
2 or 3 credits
BIOL 74501 Phytochemistry: Lecture; 30 hours lecture, 2 credits
BIOL 74502 Phytochemistry: Laboratory; 90 hours laboratory,
3 credits
BIOL 75200 Plant Morphogenesis: Lecture; 45 hours lecture,
3 credits
BIOL 75201 Plant Morphogenesis: Laboratory; 90 hours laboratory,
3 credits
BIOL 75401 Comparative Morphology of Vascular Plants: Lecture; 30
hours lecture, 2 credits
BIOL 75402 Comparative Morphology of Vascular Plants: Laboratory; 90
hours laboratory, 3 credits
BIOL 76403 Plant Ecology: Lecture; 45 hours lecture, 3 credits
BIOL 76404 Plant Ecology: Laboratory; 90 hours laboratory,
3 credits
BIOL 76405 Economic Botany; 30 hours lecture, 30 hours laboratory,
3 credits
BIOL 76501 Paleobotany: Lecture; 30 hours lecture, 2 credits
BIOL 76502 Paleobotany: Laboratory; 60 hours laboratory, 2
credits
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Seminars: General and Special Topics
BIOL 79001 Seminar in Evolution; 30 hours plus conferences,
3 credits each semester
BIOL 79002 Seminar in Genetics; 30 hours plus conferences,
3 credits each semester
BIOL 79003 Seminar in Behavioral Genetics; 30 hours plus conferences,
3 credits each semester
BIOL 79004 Seminar in Molecular Genetics; 30 hours plus conferences,
3 credits each semester
BIOL 79005 Seminar in Developmental Biology; 30 hours plus
conferences, 3 credits each semester
BIOL 79006 Seminar in Ecology; 30 hours plus conferences,
3 credits each semester
BIOL 79007 Seminar in Cytology; 45 hours, 3 credits each semester
BIOL 79008 Seminar in Biomathematics; 30 hours plus conferences,
3 credits each semester
BIOL 79009 Seminar in Biophysics; 30 hours plus conferences,
3 credits each semester
BIOL 79010 Seminar in Biochemistry; 30 hours plus conferences,
3 credits each semester
BIOL 79011 Seminar in Systematics; 45 hours, 3 credits each
semester
BIOL 79012 Seminar in Zoogeography; 45 hours, 3 credits each
semester
BIOL 79021 Seminar in Physiology; 45 hours, 3 credits each
semester
BIOL 79022 Seminar in Animal Behavior; 30 hours plus conferences,
3 credits each semester
BIOL 79023 Seminar in Cell Biology; 30 hours plus conferences,
3 credits each semester
BIOL 79063 Seminar in Biological Oceanography; 30 hours plus
conferences, 3 credits each semester
BIOL 79064 Seminar in Behavioral Aspects of Ecology; 30 hours
plus conferences, 3 credits each semester
BIOL 79065 Seminar in Tropical Forest Ecology and Conservation; 45
hours, 3 credits
BIOL 79091 Selected Topics in Animal Behavior - Biopsychology; 15
hours, 1 credit
BIOL 79093 Seminar in Acoustic Communication in Animals; 30
hours plus conferences, 3 credits each semester
BIOL 79301, 79302, 79303 Seminar in Special Topics; 15, 30,
or 45 hours, 1, 2, or 3 credits each semester
BIOL 79401 Experimental Biology: Lecture; 30 or 45 hours,
2 or 3 credits each semester
BIOL 79402 Experimental Biology: Laboratory; 60 or 90 hours
laboratory, 2 or 3 credits each semester
BIOL 79500 Basic Laboratory Techniques for Research; 15 hours
lecture, 60 hours laboratory, 3 credits each semester
BIOL 79501 Laboratory in Biotechnology; 45 hours, 3 credits
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General
BIOL 79100 Colloquium; 15 or 30 hours each semester, 1/2 or
1 credit each semester
BIOL 79200 Tutorial; 15, 30, 45, or 60 hours each semester,
1 through 4 credits each semester
BIOL 79201 Advanced Study; 15, 30, 45, or 60 hours each semester,
1-4 credits each semester
BIOL 89900 Independent Doctoral Research; Credit to be assigned,
up to a maximum of 10 credits. Required of all candidates for
the doctorate.
BIOL 90000 Dissertation Supervision; 1 credit
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